Type: MEXICO. Puebla. between 1.1 and 4.9 mi S of Zapotitlán on MEX 125 from Tehuacán to Huajuapan de León. Rocky soil on exposed, dry slopes with thorn-scrub vegetation, 5500—5700 ft, 25 Jul 1987, J. Utley & K. Utley 7948 (holotype: ♀ and ♂ MEXU; isotypes: B, BM, CAS, CU, GH, M, MICH, MO, NY, TEX, US, USF, XAL).