Type: ECUADOR: Morona-Santiago: cantón Limón Indanza, Cerro Ijiach Naint, Cordillera de Huaracayo, to the east of the river Coangos, on a sandstone mesa « tepui », 03°15´47´´S, 78°10´32´W´, 1900 m, 20 March 2001, J.M. Manzanares, D. Neill, P. Berry, L. Jost, B. Patterson and Shuar participants from Tinkimints 7213 (holotype QCNE, isotypes MO, SEL, WU).