Type: COLOMBIA. Antioquia: Municipio Salgar, 11 km on the road between Salgar and Bolombolo, very moist rocky gully, 900 m. alt., mouth of the Rio Barroso, 5° 55' north latitude, 1.X.1987, Julio Betancur, J. Zaruchi, & A. Brant 518 (HOLOTYPE: US; Isotype: HUA).