Type: Wurdack & Adderley 43601 (holotype, US; isotype, NY), terrestrial, locally abundant on shaded rocky banks of black water cano from Cerro Aracamuni on left bank of Rio Siapa just above Raudal Gallineta (about 115 river km from mouth), 130-140 m alt, Amazonas, Venezuela, 23 Jul 1959.