Type: Panama, Cocle, hills N of El Valle, E slope of Cerro Gaital, 8º.40' N, 800º 07' W, 900-1,000 m. alt., 29 May 1982, S. Knapp 5320. HOLOTYPUS: MO. PARATYPUS: ibidem, mountains around El Valle, 600-800 m alto, Feb. 1992, W Berg P40 (SEL); ibidem, vicinity of El Valle, Nov. 1993, C. Skotak s. n. (SEL).