Type: COLOMBIA: BOYACA: On damp rocky soil, cerro 1 km east southeast of Laguna Agua Clara, Serrania Pena Negra, northwest to north of Duitama, Paramo de La Rusia, 3970 m alt, 18 Dec 1972, Cleef 7430 (US, type; COL, U, isotypes); paramo with Calamagrostis and Espeletia, southeast side of Cerro Pan d'Azucar, high massif north of Belen, Municipio Tutasa, 4100 m alt, 7 May 1973, Cleef 9818 (COL, U, US).