Type: Venezuela. Amazonas: Dept. Atabapo, Cerro Marahuaca, shaded face of wet sandstone bluff, southwest-facing slopes, headwaters of Rio Iguapo, southern sector of Meseta Sur-Este, 1,560 m alt., Lat 3°36'00"N, Long. 65º23' 10"W, 13-14 October 1983, Julian A. Steyermark 129646 (holotype, US; isotype, VEN).