Type: Panama. Prov. Veraguas: Distr. Santa Fe, Santa Fe National Park, premontane rain forests, trail to Cascada Charco El Conejo close to the Quebrada Loma Grande and ANAM Station, 08°31'48.8"N, 81°08'58.8"W, at 767 m, 4 December 2012, D.A. Cáceres González 4385 (holotype: UCH; isotypes: FR and PMA).