= P. alpestris subsp. zoellneri x P. venusta
In 2006, a plant could be observed at Pichidangui (Zizka 8119 [CONC, FR]) that appeared to be intermediate between P. alpestris subsp. zoellneri and P. venusta, both occurring sympatrically in the area. The Pichidangui plant resembled the type specimen of P. berteroniana in having an inflorescence similar to P. alpestris but lacking the sterile terminal parts;
Interestingly, the further investigation of this specimen revealed it to be genetically intermediate between P. alpestris and the P. coerulea group (Schulte et al., 2010). Nonetheless, for P. coerulea does not occur in the area of Pichidangui, considering it as one of the parental species is not yet convincing.
There has been some confusion about the correct epithet of P. berteroniana. Stuessy and Marticorena (1990) proposed to change the original name to P. berteroana. However, P. berteroniana is the correct spelling according to the ICBN (McNeill et al., 2007), art. 60.6., ex. 14. —SeeZizka et al. 2013p. 399
